Stomiidae
Stomiidae is a family of deep-sea ray-finned fish, including the barbeled dragonfishes, stareaters and loosejaws.
Stomiids are generally elongated fish with black or near-black bodies, but they are highly variable in form, and are sometimes grouped into multiple different families as a result. The largest species are about 40 centimetres (16 in) in length, with most being about half that. Most species lack any scales, and have numerous small light-producing photophores scattered over their bodies. In some species, a larger photophore dangles from a barbel attached to the lower jaw, presumably as a lure to attract prey. Many species also have photophores attached to the pectoral fins, which are highly mobile, allowing the light to be moved about.[1]
They are predators, eating smaller fish, and have greatly enlarged, fang-like, teeth. Their gut is heavily pigmented, so that any luminescent organs in their prey cannot shine through their bodies and attract larger predators.[1]
Like many other deep-water fishes, stomiids produce buoyant eggs, that float up into the surface waters to hatch. The larvae live among the plankton close to the surface, and only swim down to the depths when they begin to take on adult form. Some species are known to be able to change from male to female as they age, increasing their chances of finding a mate.[1]
Species
There are 277 species in 27 genera:
- Genus Aristostomtias
- Aristostomias grimaldii Zugmayer, 1913
- Aristostomias lunifer Regan & Trewavas, 1930
- Aristostomias polydactylus Regan & Trewavas, 1930
- Shiny loosejaw, Aristostomias scintillans (Gilbert, 1915)
- Aristostomias tittmanni Welsh, 1923
- Aristostomias xenostoma Regan & Trewavas, 1930
- Genus Astronesthes
- Astronesthes atlanticus Parin & Borodulina, 1996
- Astronesthes bilobatus Parin & Borodulina, 1996
- Astronesthes boulengeri Gilchrist, 1902
- Astronesthes caulophorus Regan & Trewavas, 1929
- Astronesthes chrysophekadion (Bleeker, 1849)
- Astronesthes cyaneus (Brauer, 1902)
- Astronesthes cyclophotus Regan & Trewavas, 1929
- Astronesthes decoratus Parin & Borodulina, 2002
- Astronesthes dupliglandis Parin & Borodulina, 1997
- Astronesthes exsul Parin & Borodulina, 2002
- Astronesthes fedorovi Parin & Borodulina, 1994
- Astronesthes formosana Liao et al., 2006
- Astronesthes galapagensis Parin, Borodulina & Hulley, 1999
- Snaggletooth, Astronesthes gemmifer Goode & Bean, 1896
- Astronesthes gibbsi Borodulina, 1992
- Astronesthes gudrunae Parin & Borodulina, 2002
- Astronesthes haplophos Parin & Borodulina, 2002
- Astronesthes ijimai Tanaka, 1908
- Astronesthes illuminatus Parin, Borodulina & Hulley, 1999
- Astronesthes indicus Brauer, 1902
- Astronesthes indopacificus Parin & Borodulina, 1997
- Astronesthes karsteni Parin & Borodulina, 2002
- Astronesthes kreffti Gibbs & McKinney, 1988
- Astronesthes lamellosus Goodyear & Gibbs, 1970
- Astronesthes lampara Parin & Borodulina, 1998
- Astronesthes leucopogon Regan & Trewavas, 1929
- Astronesthes longiceps Regan & Trewavas, 1929
- Astronesthes lucibucca Parin & Borodulina, 1996
- Pacific astronesthid fish, Astronesthes lucifer Gilbert, 1905
- Astronesthes luetkeni Regan & Trewavas, 1929
- Astronesthes lupina Whitley, 1941
- Astronesthes macropogon Goodyear & Gibbs, 1970
- Astronesthes martensii Klunzinger, 1871
- Astronesthes micropogon Goodyear & Gibbs, 1970
- Astronesthes neopogon Regan & Trewavas, 1929
- Astronesthes niger Richardson, 1845
- Astronesthes nigroides Gibbs & Aron, 1960
- Astronesthes oligoa Parin & Borodulina, 2002
- Astronesthes psychrolutes (Gibbs & Weitzman, 1965)
- Astronesthes quasiindicus Parin & Borodulina, 1996
- Astronesthes richardsoni (Poey, 1852)
- Astronesthes similus Parr, 1927
- Astronesthes spatulifer Gibbs & McKinney, 1988
- Astronesthes splendida Brauer, 1902
- Astronesthes tanibe Parin & Borodulina, 2001
- Astronesthes tatyanae Parin & Borodulina, 1998
- Astronesthes tchuvasovi Parin & Borodulina, 1996
- Astronesthes trifibulatus Gibbs, Amaoka & Haruta, 1984
- Astronesthes zetgibbsi Parin & Borodulina, 1997
- Astronesthes zharovi Parin & Borodulina, 1998
- Genus Bathophilus
- Bathophilus abarbatus Barnett & Gibbs, 1968
- Bathophilus altipinnis Beebe, 1933
- Bathophilus ater (Brauer, 1902)
- Bathophilus brevis Regan & Trewavas, 1930
- Bathophilus digitatus (Welsh, 1923)
- Sparing, Bathophilus filifer (Regan & Trewavas, 1929)
- Highfin dragonfish, Bathophilus flemingi Aron & McCrery, 1958
- Bathophilus indicus (Brauer, 1902)
- Bathophilus irregularis Norman, 1930
- Bathophilus kingi Barnett & Gibbs, 1968
- Bathophilus longipinnis (Pappenheim, 1914)
- Scaleless dragonfish, Bathophilus nigerrimus Giglioli, 1882
- Bathophilus pawneei Parr, 1927
- Bathophilus proximus Regan & Trewavas, 1930
- Bathophilus schizochirus Regan & Trewavas, 1930
- Bathophilus vaillanti (Zugmayer, 1911)
- Genus Borostomias
- Borostomias abyssorum (Koehler, 1896)
- Borostomias antarcticus (Lönnberg, 1905)
- Borostomias elucens (Brauer, 1906)
- Borostomias mononema (Regan & Trewavas, 1929)
- Borostomias pacificus (Imai, 1941)
- Panama snaggletooth, Borostomias panamensis Regan & Trewavas, 1929
- Genus Chauliodus
- Chauliodus barbatus (Lowe, 1843)
- Dana viperfish, Chauliodus danae Regan & Trewavas, 1929
- Chauliodus dentatus Garman, 1899
- Pacific viperfish, Chauliodus macouni Bean, 1890
- Chauliodus minimus Parin & Novikova, 1974
- Chauliodus pammelas Alcock, 1892
- Chauliodus schmidti (Regan & Trewavas, 1929)
- Sloane's viperfish, Chauliodus sloani Bloch & Schneider, 1801
- Chauliodus vasnetzovi Novikova, 1972
- Genus Chirostomias
- Chirostomias pliopterus Regan & Trewavas, 1930
- Genus Echiostoma
- Echiostoma barbatum Lowe, 1843
- Genus Eupogonesthes
- Eupogonesthes xenicus Parin & Borodulina, 1993
- Genus Eustomias
- Eustomias achirus Parin & Pokhil'skaya, 1974
- Eustomias acinosus Regan & Trewavas, 1930
- Eustomias aequatorialis Clarke, 1998
- Eustomias albibulbus Clarke, 2001
- Eustomias appositus Gibbs, Clarke & Gomon, 1983
- Eustomias arborifer Parr, 1927
- Eustomias australensis Gibbs, Clarke & Gomon, 1983
- Eustomias austratlanticus Gibbs, Clarke & Gomon, 1983
- Eustomias bertelseni Gibbs, Clarke & Gomon, 1983
- Eustomias bibulboides Gibbs, Clarke & Gomon, 1983
- Eustomias bibulbosus Parr, 1927
- Eustomias bifilis (Regan & Trewavas, 1929)
- Eustomias bigelowi Welsh, 1923
- Eustomias bimargaritatus Regan & Trewavas, 1930
- Eustomias bimargaritoides Gibbs, Clarke & Gomon, 1983
- Eustomias binghami Parr, 1927
- Eustomias bituberatus Regan & Trewavas, 1930
- Eustomias bituberoides Gibbs, Clarke & Gomon, 1983
- Eustomias borealis Clarke, 2000
- Eustomias braueri (Regan, 1908)
- Eustomias brevibarbatus Parr, 1927
- Eustomias bulbiramis Clarke, 2001
- Eustomias bulbornatus Gibbs, 1960
- Eustomias cancriensis Gibbs, Clarke & Gomon, 1983
- Eustomias cirritus Gibbs, Clarke & Gomon, 1983
- Eustomias contiguus Gomon & Gibbs, 1985
- Eustomias crossotus Gibbs, Clarke & Gomon, 1983
- Eustomias crucis Gibbs & Craddock, 1973
- Eustomias cryptobulbus Clarke, 2001
- Eustomias curtatus Gibbs, Clarke & Gomon, 1983
- Eustomias curtifilis Clarke, 2000
- Eustomias danae (Regan & Trewavas, 1929)
- Eustomias decoratus (Parin & Borodulina, 2002)
- Eustomias dendriticus Regan & Trewavas, 1930
- Eustomias deofamiliaris Gibbs, Clarke & Gomon, 1983
- Eustomias digitatus (Welsh, 1923)
- Eustomias dinema Clarke, 1999
- Eustomias dispar Gomon & Gibbs, 1985
- Eustomias dubius Parr, 1927
- Eustomias elongatus Clarke, 2001
- Eustomias enbarbatus Welsh, 1923
- Eustomias filifer (Regan & Trewavas, 1929)
- Eustomias fissibarbis (Pappenheim, 1914)
- Eustomias flagellifer Clarke, 2001
- Eustomias furcifer Regan & Trewavas, 1930
- Eustomias gibbsi (Borodulina, 1992)
- Eustomias grandibulbus Gibbs, Clarke & Gomon, 1983
- Eustomias hulleyi Gomon & Gibbs, 1985
- Eustomias hypopsilus Gomon & Gibbs, 1985
- Eustomias ignotus Gomon & Gibbs, 1985
- Eustomias inconstans Gibbs, Clarke & Gomon, 1983
- Eustomias insularum Clarke, 1998
- Eustomias intermedius Clarke, 1998
- Eustomias interruptus Clarke, 1999
- Eustomias ioani Parin & Pokhil'skaya, 1974
- Eustomias jimcraddocki Sutton & Hartel, 2004
- Eustomias kreffti (Gibbs & McKinney, 1988)
- Eustomias lanceolatus Clarke, 1999
- Eustomias leptobolus Regan & Trewavas, 1930
- Eustomias lipochirus Regan & Trewavas, 1930
- Eustomias longibarba Parr, 1927
- Eustomias longiramis Clarke, 2001
- Eustomias macronema Regan & Trewavas, 1930
- Eustomias macrophthalmus (Regan & Trewavas, 1929)
- Eustomias macrurus Regan & Trewavas, 1930
- Eustomias magnificus Clarke, 2001
- Eustomias medusa Gibbs, Clarke & Gomon, 1983
- Eustomias melanonema Regan & Trewavas, 1930
- Eustomias melanostigma Regan & Trewavas, 1930
- Eustomias melanostigmoides Gibbs, Clarke & Gomon, 1983
- Eustomias mesostenus Gibbs, Clarke & Gomon, 1983
- Eustomias metamelas Gomon & Gibbs, 1985
- Eustomias micraster Parr, 1927
- Eustomias micropterygius Parr, 1927
- Eustomias minimus (Parin & Novikova, 1974)
- Eustomias monoclonoides Clarke, 1999
- Eustomias monoclonus Regan & Trewavas, 1930
- Eustomias monodactylus Regan & Trewavas, 1930
- Eustomias multifilis Parin & Pokhil'skaya, 1978
- Eustomias obscurus Vaillant, 1884
- Eustomias orientalis Gibbs, Clarke & Gomon, 1983
- Eustomias pacificus (Imai, 1941)
- Eustomias parini Clarke, 2001
- Eustomias parri Regan & Trewavas, 1930
- Eustomias patulus Regan & Trewavas, 1930
- Eustomias paucifilis Parr, 1927
- Eustomias paxtoni Clarke, 2001
- Eustomias perplexus Gibbs, Clarke & Gomon, 1983
- Eustomias pinnatus Clarke, 1999
- Eustomias polyaster Parr, 1927
- Eustomias posti Gibbs, Clarke & Gomon, 1983
- Eustomias precarius Gomon & Gibbs, 1985
- Eustomias problematicus Clarke, 2001
- Eustomias pyrifer Regan & Trewavas, 1930
- Eustomias quadrifilis Gomon & Gibbs, 1985
- Eustomias radicifilis Borodin, 1930
- Eustomias satterleei Beebe, 1933
- Eustomias schiffi Beebe, 1932
- Eustomias schmidti (Regan & Trewavas, 1929)
- Eustomias silvescens Regan & Trewavas, 1930
- Eustomias similis (Parr, 1927)
- Eustomias simplex (Parr, 1927)
- Eustomias spherulifer Gibbs, Clarke & Gomon, 1983
- Eustomias suluensis Gibbs, Clarke & Gomon, 1983
- Eustomias tenisoni Regan & Trewavas, 1930
- Eustomias tetranema Zugmayer, 1913
- Eustomias teuthidopsis Gibbs, Clarke & Gomon, 1983
- Eustomias tomentosis Clarke, 1998
- Eustomias trewavasae Norman, 1930
- Eustomias triramis Regan & Trewavas, 1930
- Eustomias uniramis Clarke, 1999
- Eustomias variabilis Regan & Trewavas, 1930
- Eustomias vitiazi Parin & Pokhil'skaya, 1974
- Eustomias vulgaris Clarke, 2001
- Eustomias woollardi Clarke, 1998
- Eustomias xenobolus Regan & Trewavas, 1930
- Genus Flagellostomias
- Flagellostomias boureei (Zugmayer, 1913)
- Genus Grammatostomias
- Grammatostomias circularis Morrow, 1959
- Grammatostomias dentatus (Garman, 1899)
- Grammatostomias flagellibarba Holt & Byrne, 1910
- Genus Heterophotus
- Heterophotus ophistoma Regan & Trewavas, 1929
- Genus Idiacanthus
- Genus Leptostomias
- Leptostomias analis Regan & Trewavas, 1930
- Leptostomias bermudensis Beebe, 1932
- Leptostomias bilobatus (Parin & Borodulina, 1996)
- Leptostomias gladiator (Zugmayer, 1911)
- Leptostomias gracilis Regan & Trewavas, 1930
- Leptostomias haplocaulus Regan & Trewavas, 1930
- Leptostomias leptobolus (Regan & Trewavas, 1930)
- Leptostomias longibarba (Parr, 1927)
- Long-threaded dragonfish, Leptostomias macronema (Regan & Trewavas, 1930)
- Leptostomias macropogon (Goodyear & Gibbs, 1970)
- Leptostomias multifilis (Parin & Pokhil'skaya, 1978)
- Leptostomias robustus Imai, 1941
- Genus Malacosteus
- Genus Melanostomias
- Melanostomias bartonbeani Parr, 1927
- Melanostomias biseriatus Regan & Trewavas, 1930
- Melanostomias globulifer Fowler, 1934
- Melanostomias macrophotus Regan & Trewavas, 1930
- Melanostomias margaritifer Regan & Trewavas, 1930
- Melanostomias melanopogon Regan & Trewavas, 1930
- Melanostomias melanops Brauer, 1902
- Melanostomias niger (Richardson, 1845)
- Melanostomias nigroaxialis Parin & Pokhil'skaya, 1978
- Melanostomias paucilaternatus Parin & Pokhil'skaya, 1978
- Melanostomias pauciradius Matsubara, 1938
- Melanostomias pollicifer Parin & Pokhil'skaya, 1978
- Melanostomias stewarti Fowler, 1934
- Melanostomias tentaculatus (Regan & Trewavas, 1930)
- Valdivia black dragonfish, Melanostomias valdiviae Brauer, 1902
- Melanostomias vierecki Fowler, 1934
- Genus Neonesthes
- Genus Odontostomias
- Odontostomias masticopogon Norman, 1930
- Odontostomias micropogon (Goodyear & Gibbs, 1970)
- Genus Opostomias
- Opostomias micripnus (Günther, 1878)
- Pitgum lanternfish, Opostomias mitsuii Imai, 1941
- Genus Pachystomias
- Pachystomias microdon (Günther, 1878)
- Genus Photonectes
- Photonectes achirus (Parin & Pokhil'skaya, 1974)
- Photonectes albipennis (Döderlein, 1882)
- Photonectes barnetti Klepadlo, 2011[2]
- Photonectes braueri (Regan, 1908)
- Photonectes caerulescens Regan & Trewavas, 1930
- Photonectes coffea Klepadlo, 2011[2]
- Photonectes corynodes Klepadlo, 2011[2]
- Photonectes dinema (Clarke, 1999)
- Photonectes gracilis (Regan & Trewavas, 1930)
- Photonectes leucospilus Regan & Trewavas, 1930
- Photonectes margarita (Goode & Bean, 1896)
- Photonectes mirabilis Parr, 1927
- Photonectes munificus Gibbs, 1968
- Photonectes parvimanus Regan & Trewavas, 1930
- Photonectes phyllopogon Regan & Trewavas, 1930
- Genus Photostomias
-
- Photostomias guernei Collett, 1889
- Photostomias mirabilis (Parr, 1927)
- Genus Rhadinesthes
- Rhadinesthes decimus (Zugmayer, 1911)
- Genus Stomias
- Günther's boafish, Stomias affinis Günther, 1887
- Black-belly dragonfish, Stomias atriventer Garman, 1899
- Scaly dragonfish, Stomias boa boa (Risso, 1810)
- Stomias boa colubrinus Garman, 1899
- Boa dragonfish, Stomias boa ferox Reinhardt, 1842
- Stomias brevibarbatus (Parr, 1927)
- Stomias danae (Regan & Trewavas, 1929)
- Stomias gracilis (Regan & Trewavas, 1930)
- Stomias lampropeltis Gibbs, 1969
- Stomias longibarbatus (Brauer, 1902)
- Alcock's boafish, Stomias nebulosus Alcock, 1889
- Genus Tactostoma
- Longfin dragonfish, Tactostoma macropus Bolin, 1939
- Genus Thysanactis
- Thysanactis dentex Regan & Trewavas, 1930
- Genus Trigonolampa
- Threelight dragonfish, Trigonolampa miriceps Regan & Trewavas, 1930
References
- ^ a b c Fink, William L. (1998). Paxton, J.R. & Eschmeyer, W.N.. ed. Encyclopedia of Fishes. San Diego: Academic Press. p. 122. ISBN 0-12-547665-5.
- ^ a b c Klepadlo, C. (2011). "Three New Species of the Genus Photonectes (Teleostei: Stomiiformes: Stomiidae: Melanostomiinae) from the Pacific Ocean." Copeia 2011.2: 201-210.